This image of Grizzly 399 was taken when her quadruplets were still cubs-of-the-year. Grand Teton National Park’s most famous grizzly bear, known by her research number 399, has raised litter after litter along the Oxbow Bend of the Snake River. A marvel of nature, thousands of visitors from around the world watch 399 and her offspring in and near the park. She is regarded as an ambassador for all grizzlies because of her lifelong public profile and the educational opportunities she provides.